April 9 - 21, 2015
Application Deadline: February 23, 2015

Mustafa Khokha,
Yale University
Karen Liu, King's College London, UK

See the roll of honor - who's taken the course in the past

Techniques covered in this course will include microinjection, molecular manipulations such as CRISPR knockouts, transgenics, and mRNA overexpression. These can be combined with explant and transplant methods to simplify or test tissue level interactions. To visualize subcellular and intercellular activities, we will introduce a variety of imaging methods including time-lapse, fluorescent and confocal microscopy. Additional methods include mRNA in situ hybridization and protein immunohistochemistry as well as basic bioinformatic techniques for gene comparison and functional analysis.

This course is designed for those new to the Xenopus field, as well as for more advanced students who are interested in emerging technologies. We encourage students to bring their own genes of interest and will tailor aspects of the course to enable them to initiate studies on their specific projects.

2015 Speakers include:
Helene Cousin, University of Massachusetts, Amherst
Heithem El-Hodiri, Ohio State University
Scott Fraser, University of Southern California
Rebecca Heald, University of California, Berkeley
Raymond Keller, University of Virginia
Marc Kirschner, Harvard Medical School
Rachel Miller, UT Medical School at Houston
Nancy Papalopulu, University of Manchester, UK
John Wallingford, University of Texas at Austin

This course is supported with funds provided by: National Institute of Child Health and Human Development and Howard Hughes Medical Institute

Cost (including tuition, board and lodging): $3,735

This button links to a short form which confirms your interest in the course.
No fees are due until you have completed the full application process and
are accepted into the course.


Students accepted into the course should plan to arrive by early evening on April 8 and plan to depart after lunch on April 21.